| Andrew William Oliver |
205,882,352,941 |
£288,235,294 |
| TD Direct Investing Nominees |
94,495,144,706 |
£132,293,203 |
| HSDL Nominees Limited |
81,968,497,647 |
£114,755,897 |
| David Lenigas |
79,411,764,706 |
£111,176,471 |
| Greenhair Services Limited |
73,235,294,118 |
£102,529,412 |
| Academy Minerals Limited |
70,588,235,294 |
£98,823,529 |
| Kavi Dhana |
44,021,067,059 |
£61,629,494 |
| IP2IPO Limited (together with Techtran Group Limited) |
34,418,058,824 |
£48,185,282 |
| XCAP Nominees Limited |
19,824,847,059 |
£27,754,786 |
| IP Venture Fund Limited |
16,302,105,882 |
£22,822,948 |
| Mr Mark V Houghton-Berry |
12,335,376,471 |
£17,269,527 |
| The University of Leeds |
9,552,264,706 |
£13,373,171 |
| The Wellcome Trust Limited |
6,107,858,824 |
£8,551,002 |
| Adrian Stiff |
37,000,000 |
£51,800 |
| Hargreaves Lansdown |
27,752,212 |
£38,853 |
| JIM Nominees Limited |
27,116,707 |
£37,963 |
| Pershing Nominees |
26,006,789 |
£36,410 |
| Hargreaves Lansdown (Nominees) Limited Des 15942 |
25,609,943 |
£35,854 |
| Chris Akers |
25,000,000 |
£35,000 |
| Vidacos Nominees Limited |
20,709,150 |
£28,993 |
| The Bank OF New York (Nominees) Limited Des 672938 |
19,705,371 |
£27,588 |
| Oberon Investments |
19,705,371 |
£27,588 |
| Barclays Direct Investing Nominees Limited |
19,358,393 |
£27,102 |
| Winterflood Securities Limited |
18,278,266 |
£25,590 |
| Donald Strang |
17,820,211 |
£24,948 |
| Interactive Investor Services Nominees Limited |
17,219,649 |
£24,108 |
| ABN Amro Bank |
13,801,956 |
£19,323 |
| Hargreaves Lansdown (Nominees) Limited Des VRA |
12,572,701 |
£17,602 |
| David Graham Brown |
12,100,000 |
£16,940 |
| Sebastian Marr |
8,180,672 |
£11,453 |