| MassMutual Life Insurance Company |
12,727,105 |
£523,338,558 |
| MFS Investment Management |
11,499,077 |
£472,842,046 |
| Fidelity Management & Research Company |
8,063,819 |
£331,584,237 |
| MI Metron UK Bidco Limited |
7,691,714 |
£316,283,280 |
| Societe Generale |
7,177,114 |
£295,122,928 |
| Standard Life Investment Ltd |
5,920,593 |
£243,454,784 |
| OppenheimerFunds, Inc. |
5,720,217 |
£235,215,323 |
| Marathon Asset Management LLP |
5,708,763 |
£234,744,335 |
| Vanguard Group |
5,394,997 |
£221,842,277 |
| UBS Asset Management |
5,231,636 |
£215,124,872 |
| F & C Asset Management plc |
5,212,550 |
£214,340,056 |
| Massachusetts Financial Services Company |
5,178,500 |
£212,939,920 |
| JPMorgan Chase & Co. |
4,841,669 |
£199,089,429 |
| Wellington Management Company |
4,248,835 |
£174,712,095 |
| Sprucegrove Investment Management |
3,832,503 |
£157,592,523 |
| Artemis Investment Management |
3,661,077 |
£150,543,486 |
| Royal London Asset Management Limited |
3,626,169 |
£149,108,069 |
| Schroder Investment Management |
3,484,265 |
£143,272,977 |
| BlackRock, Inc. |
3,429,918 |
£141,038,228 |
| Ninety One |
3,160,918 |
£129,976,948 |
| Liontrust Asset Management |
3,052,149 |
£125,504,367 |
| Evenlode Investment |
2,971,067 |
£122,170,275 |
| Ignis Investment Services Limited |
10 |
£411 |
| Ameriprise Financial Inc. |
1 |
£41 |
| COLUMBIA WANGER ASSET MANAGEMENT, L.P. |
1 |
£41 |