| Gresham House Asset Management Ltd |
3,746,521 |
£60,131,662 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£52,621,562 |
| Liontrust Investment Services Limited |
2,457,011 |
£39,435,027 |
| Octopus Investments Nominees Limited |
2,335,294 |
£37,481,469 |
| Aberforth Partners LLP |
2,067,205 |
£33,178,640 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£17,847,841 |
| Brooks Macdonald Asset Management |
1,105,946 |
£17,750,433 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,391,672 |
| Jupiter Fund Management plc |
938,000 |
£15,054,900 |
| Invesco Asset Management |
817,151 |
£13,115,274 |
| Invesco |
817,151 |
£13,115,274 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£12,987,660 |
| The Wellcome Trust Limited |
805,370 |
£12,926,188 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,622,555 |
| Hargreave Hale & Co |
784,942 |
£12,598,319 |
| Canaccord Genuity Group Inc |
780,008 |
£12,519,128 |
| Artemis Investment Management Ltd |
729,575 |
£11,709,679 |
| Norges Bank |
683,030 |
£10,962,632 |
| Charles Stanley |
659,277 |
£10,581,396 |
| Jonathan M Gumpel |
642,460 |
£10,311,483 |
| Brewin Dolphin Limited |
622,157 |
£9,985,620 |
| Chelverton Asset Management |
609,008 |
£9,774,578 |
| JTC plc |
603,261 |
£9,682,339 |
| Amati Global Investors |
572,459 |
£9,187,967 |
| Rathbones Investment Management |
488,191 |
£7,835,466 |
| Kames Capital |
487,376 |
£7,822,385 |
| Spearpoint Holdings Limited |
428,227 |
£6,873,043 |