| Gresham House Asset Management Ltd |
3,884,684 |
£52,249,000 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£44,097,197 |
| Octopus Investments Nominees Limited |
2,335,294 |
£31,409,704 |
| Liontrust Investment Services Limited |
1,743,782 |
£23,453,868 |
| Aberforth Partners LLP |
1,606,852 |
£21,612,159 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£14,956,602 |
| Brooks Macdonald Asset Management |
1,105,946 |
£14,874,974 |
| Canaccord Genuity Group Inc |
1,021,288 |
£13,736,324 |
| Jupiter Fund Management plc |
938,000 |
£12,616,100 |
| Invesco Asset Management |
817,151 |
£10,990,681 |
| Invesco |
817,151 |
£10,990,681 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£10,883,740 |
| The Wellcome Trust Limited |
805,370 |
£10,832,226 |
| FIDELITY INTERNATIONAL |
786,452 |
£10,577,779 |
| Hargreave Hale & Co |
784,942 |
£10,557,470 |
| Canaccord Genuity Group Inc |
780,008 |
£10,491,108 |
| Artemis Investment Management Ltd |
729,575 |
£9,812,784 |
| Norges Bank |
683,030 |
£9,186,754 |
| Charles Stanley |
659,277 |
£8,867,276 |
| Jonathan M Gumpel |
642,460 |
£8,641,087 |
| Brewin Dolphin Limited |
622,157 |
£8,368,012 |
| Chelverton Asset Management |
609,008 |
£8,191,158 |
| JTC plc |
603,261 |
£8,113,860 |
| Amati Global Investors |
572,459 |
£7,699,574 |
| Rathbones Investment Management |
488,191 |
£6,566,169 |
| Kames Capital |
487,376 |
£6,555,207 |
| Spearpoint Holdings Limited |
428,227 |
£5,759,653 |