| Gresham House Asset Management Ltd |
3,746,521 |
£60,880,966 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£53,277,282 |
| Liontrust Investment Services Limited |
2,457,011 |
£39,926,429 |
| Octopus Investments Nominees Limited |
2,335,294 |
£37,948,528 |
| Brooks Macdonald Asset Management |
1,105,946 |
£17,971,622 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,595,930 |
| Jupiter Fund Management plc |
938,000 |
£15,242,500 |
| Aberforth Partners LLP |
898,030 |
£14,592,988 |
| Invesco Asset Management |
817,151 |
£13,278,704 |
| Invesco |
817,151 |
£13,278,704 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£13,149,500 |
| The Wellcome Trust Limited |
805,370 |
£13,087,262 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,779,845 |
| Hargreave Hale & Co |
784,942 |
£12,755,308 |
| Canaccord Genuity Group Inc |
780,008 |
£12,675,130 |
| Artemis Investment Management Ltd |
729,575 |
£11,855,594 |
| Norges Bank |
683,030 |
£11,099,238 |
| Charles Stanley |
659,277 |
£10,713,251 |
| Jonathan M Gumpel |
642,460 |
£10,439,975 |
| Brewin Dolphin Limited |
622,157 |
£10,110,051 |
| JTC plc |
603,261 |
£9,802,991 |
| Amati Global Investors |
572,459 |
£9,302,459 |
| Chelverton Asset Management |
569,750 |
£9,258,438 |
| Brooks Macdonald Employee Benefit Trust |
552,633 |
£8,980,286 |
| Rathbones Investment Management |
488,191 |
£7,933,104 |
| Kames Capital |
487,376 |
£7,919,860 |
| Spearpoint Holdings Limited |
428,227 |
£6,958,689 |