| Alantra EQMC Asset Management SGIIC, S.A. |
4,876,250 |
£94,794,300 |
| EQMC Europe Development Capital Fund |
4,858,350 |
£94,446,324 |
| Kempen Capital Management N.V. |
3,328,822 |
£64,712,300 |
| BlackRock Smaller Companies Trust Plc |
3,206,055 |
£62,325,709 |
| Capital Research and Management Company |
2,990,645 |
£58,138,139 |
| Aberdeen Asset Management plc |
1,835,099 |
£35,674,325 |
| Franklin Templeton Investments |
1,700,000 |
£33,048,000 |
| Threadneedle Asset Management |
1,597,700 |
£31,059,288 |
| Aggregate of Standard Life Aberdeen plc |
1,580,030 |
£30,715,783 |
| River & Mercantile Asset Management LLP |
1,549,478 |
£30,121,852 |
| Hargreave Hale & Co |
1,546,771 |
£30,069,228 |
| Cazenove Capital Management Limited |
1,538,403 |
£29,906,554 |
| Ameriprise Financial, Inc. and its group |
1,538,156 |
£29,901,753 |
| Canaccord Genuity Group Inc |
1,529,075 |
£29,725,218 |
| Polar Capital LLP |
1,517,909 |
£29,508,151 |
| Ancora Alternatives LLC |
1,504,769 |
£29,252,709 |
| MI Chelverton UK Equity Growth Fund |
1,500,000 |
£29,160,000 |
| ClearBridge Investment Management Limited |
1,486,511 |
£28,897,774 |
| Schroders plc |
1,478,291 |
£28,737,977 |
| The Capital Group Companies, Inc. |
1,443,660 |
£28,064,750 |
| BlackRock, Inc. |
1,443,454 |
£28,060,746 |
| Aberforth Partners LLP |
1,416,081 |
£27,528,615 |
| Janus Henderson Investors |
1,408,457 |
£27,380,404 |
| Wasatch Advisors, Inc. |
1,285,950 |
£24,998,868 |
| JPMorgan Asset Management Holdings Inc. |
1,256,443 |
£24,425,252 |
| JTC Share Plan Trustees (Guernsey) Limited |
878,463 |
£17,077,321 |