| Gresham House Asset Management Ltd |
3,746,521 |
£61,068,292 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£53,441,213 |
| Liontrust Investment Services Limited |
2,457,011 |
£40,049,279 |
| Octopus Investments Nominees Limited |
2,335,294 |
£38,065,292 |
| Aberforth Partners LLP |
2,067,205 |
£33,695,442 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£18,125,844 |
| Brooks Macdonald Asset Management |
1,105,946 |
£18,026,920 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,646,994 |
| Jupiter Fund Management plc |
938,000 |
£15,289,400 |
| Invesco Asset Management |
817,151 |
£13,319,561 |
| Invesco |
817,151 |
£13,319,561 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£13,189,960 |
| The Wellcome Trust Limited |
805,370 |
£13,127,531 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,819,168 |
| Hargreave Hale & Co |
784,942 |
£12,794,555 |
| Canaccord Genuity Group Inc |
780,008 |
£12,714,130 |
| Artemis Investment Management Ltd |
729,575 |
£11,892,072 |
| Norges Bank |
683,030 |
£11,133,389 |
| Charles Stanley |
659,277 |
£10,746,215 |
| Jonathan M Gumpel |
642,460 |
£10,472,098 |
| Brewin Dolphin Limited |
622,157 |
£10,141,159 |
| Chelverton Asset Management |
609,008 |
£9,926,830 |
| JTC plc |
603,261 |
£9,833,154 |
| Amati Global Investors |
572,459 |
£9,331,082 |
| Rathbones Investment Management |
488,191 |
£7,957,513 |
| Kames Capital |
487,376 |
£7,944,229 |
| Spearpoint Holdings Limited |
428,227 |
£6,980,100 |