| Andrew William Oliver |
205,882,352,941 |
£334,558,824 |
| TD Direct Investing Nominees |
94,495,144,706 |
£153,554,610 |
| HSDL Nominees Limited |
81,968,497,647 |
£133,198,809 |
| David Lenigas |
79,411,764,706 |
£129,044,118 |
| Greenhair Services Limited |
73,235,294,118 |
£119,007,353 |
| Academy Minerals Limited |
70,588,235,294 |
£114,705,882 |
| Kavi Dhana |
44,021,067,059 |
£71,534,234 |
| IP2IPO Limited (together with Techtran Group Limited) |
34,418,058,824 |
£55,929,346 |
| XCAP Nominees Limited |
19,824,847,059 |
£32,215,376 |
| IP Venture Fund Limited |
16,302,105,882 |
£26,490,922 |
| Mr Mark V Houghton-Berry |
12,335,376,471 |
£20,044,987 |
| The University of Leeds |
9,552,264,706 |
£15,522,430 |
| The Wellcome Trust Limited |
6,107,858,824 |
£9,925,271 |
| Adrian Stiff |
37,000,000 |
£60,125 |
| Hargreaves Lansdown |
27,752,212 |
£45,097 |
| JIM Nominees Limited |
27,116,707 |
£44,065 |
| Pershing Nominees |
26,006,789 |
£42,261 |
| Hargreaves Lansdown (Nominees) Limited Des 15942 |
25,609,943 |
£41,616 |
| Chris Akers |
25,000,000 |
£40,625 |
| Vidacos Nominees Limited |
20,709,150 |
£33,652 |
| The Bank OF New York (Nominees) Limited Des 672938 |
19,705,371 |
£32,021 |
| Oberon Investments |
19,705,371 |
£32,021 |
| Barclays Direct Investing Nominees Limited |
19,358,393 |
£31,457 |
| Winterflood Securities Limited |
18,278,266 |
£29,702 |
| Donald Strang |
17,820,211 |
£28,958 |
| Interactive Investor Services Nominees Limited |
17,219,649 |
£27,982 |
| ABN Amro Bank |
13,801,956 |
£22,428 |
| Hargreaves Lansdown (Nominees) Limited Des VRA |
12,572,701 |
£20,431 |
| David Graham Brown |
12,100,000 |
£19,662 |
| Sebastian Marr |
8,180,672 |
£13,294 |