| Gresham House Asset Management Ltd |
3,746,521 |
£62,004,923 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£54,260,863 |
| Liontrust Investment Services Limited |
2,348,411 |
£38,866,202 |
| Octopus Investments Nominees Limited |
2,335,294 |
£38,649,116 |
| Aberforth Partners LLP |
1,606,852 |
£26,593,401 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£18,403,848 |
| Brooks Macdonald Asset Management |
1,105,946 |
£18,303,406 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,902,316 |
| Jupiter Fund Management plc |
938,000 |
£15,523,900 |
| Invesco Asset Management |
817,151 |
£13,523,849 |
| Invesco |
817,151 |
£13,523,849 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£13,392,260 |
| The Wellcome Trust Limited |
805,370 |
£13,328,874 |
| FIDELITY INTERNATIONAL |
786,452 |
£13,015,781 |
| Hargreave Hale & Co |
784,942 |
£12,990,790 |
| Canaccord Genuity Group Inc |
780,008 |
£12,909,132 |
| Artemis Investment Management Ltd |
729,575 |
£12,074,466 |
| Norges Bank |
683,030 |
£11,304,146 |
| Charles Stanley |
659,277 |
£10,911,034 |
| Jonathan M Gumpel |
642,460 |
£10,632,713 |
| Brewin Dolphin Limited |
622,157 |
£10,296,698 |
| Chelverton Asset Management |
609,008 |
£10,079,082 |
| JTC plc |
603,261 |
£9,983,970 |
| Amati Global Investors |
572,459 |
£9,474,196 |
| Rathbones Investment Management |
488,191 |
£8,079,561 |
| Kames Capital |
487,376 |
£8,066,073 |
| Spearpoint Holdings Limited |
428,227 |
£7,087,157 |