| Gresham House Asset Management Ltd |
3,746,521 |
£61,255,618 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£53,605,143 |
| Liontrust Investment Services Limited |
2,348,411 |
£38,396,520 |
| Octopus Investments Nominees Limited |
2,335,294 |
£38,182,057 |
| Aberforth Partners LLP |
1,606,852 |
£26,272,030 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£18,181,445 |
| Brooks Macdonald Asset Management |
1,105,946 |
£18,082,217 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,698,059 |
| Jupiter Fund Management plc |
938,000 |
£15,336,300 |
| Invesco Asset Management |
817,151 |
£13,360,419 |
| Invesco |
817,151 |
£13,360,419 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£13,230,420 |
| The Wellcome Trust Limited |
805,370 |
£13,167,800 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,858,490 |
| Hargreave Hale & Co |
784,942 |
£12,833,802 |
| Canaccord Genuity Group Inc |
780,008 |
£12,753,131 |
| Artemis Investment Management Ltd |
729,575 |
£11,928,551 |
| Norges Bank |
683,030 |
£11,167,540 |
| Charles Stanley |
659,277 |
£10,779,179 |
| Jonathan M Gumpel |
642,460 |
£10,504,221 |
| Brewin Dolphin Limited |
622,157 |
£10,172,267 |
| Chelverton Asset Management |
609,008 |
£9,957,281 |
| JTC plc |
603,261 |
£9,863,317 |
| Amati Global Investors |
572,459 |
£9,359,705 |
| Rathbones Investment Management |
488,191 |
£7,981,923 |
| Kames Capital |
487,376 |
£7,968,598 |
| Spearpoint Holdings Limited |
428,227 |
£7,001,512 |