| Gresham House Asset Management Ltd |
3,884,684 |
£54,579,810 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£46,064,358 |
| Octopus Investments Nominees Limited |
2,335,294 |
£32,810,881 |
| Liontrust Investment Services Limited |
1,743,782 |
£24,500,137 |
| Aberforth Partners LLP |
1,606,852 |
£22,576,271 |
| Liontrust Investment Partners LLP |
1,549,398 |
£21,769,042 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£15,623,811 |
| Brooks Macdonald Asset Management |
1,105,946 |
£15,538,541 |
| Canaccord Genuity Group Inc |
1,021,288 |
£14,349,096 |
| Jupiter Fund Management plc |
938,000 |
£13,178,900 |
| Invesco Asset Management |
817,151 |
£11,480,972 |
| Invesco |
817,151 |
£11,480,972 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£11,369,260 |
| The Wellcome Trust Limited |
805,370 |
£11,315,448 |
| FIDELITY INTERNATIONAL |
786,452 |
£11,049,651 |
| Hargreave Hale & Co |
784,942 |
£11,028,435 |
| Canaccord Genuity Group Inc |
780,008 |
£10,959,112 |
| Artemis Investment Management Ltd |
729,575 |
£10,250,529 |
| Norges Bank |
683,030 |
£9,596,572 |
| Charles Stanley |
659,277 |
£9,262,842 |
| Jonathan M Gumpel |
642,460 |
£9,026,563 |
| Brewin Dolphin Limited |
622,157 |
£8,741,306 |
| Chelverton Asset Management |
609,008 |
£8,556,562 |
| JTC plc |
603,261 |
£8,475,817 |
| Amati Global Investors |
572,459 |
£8,043,049 |
| Rathbones Investment Management |
488,191 |
£6,859,084 |
| Kames Capital |
487,376 |
£6,847,633 |
| Spearpoint Holdings Limited |
428,227 |
£6,016,589 |