| Gresham House Asset Management Ltd |
3,746,521 |
£59,195,032 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£51,801,912 |
| Liontrust Investment Services Limited |
2,457,011 |
£38,820,774 |
| Octopus Investments Nominees Limited |
2,335,294 |
£36,897,645 |
| Brooks Macdonald Asset Management |
1,105,946 |
£17,473,947 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,136,350 |
| Jupiter Fund Management plc |
938,000 |
£14,820,400 |
| Aberforth Partners LLP |
898,030 |
£14,188,874 |
| Invesco Asset Management |
817,151 |
£12,910,986 |
| Invesco |
817,151 |
£12,910,986 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£12,785,360 |
| The Wellcome Trust Limited |
805,370 |
£12,724,846 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,425,942 |
| Hargreave Hale & Co |
784,942 |
£12,402,084 |
| Canaccord Genuity Group Inc |
780,008 |
£12,324,126 |
| Artemis Investment Management Ltd |
729,575 |
£11,527,285 |
| Norges Bank |
683,030 |
£10,791,874 |
| Charles Stanley |
659,277 |
£10,416,577 |
| Jonathan M Gumpel |
642,460 |
£10,150,868 |
| Brewin Dolphin Limited |
622,157 |
£9,830,081 |
| JTC plc |
603,261 |
£9,531,524 |
| Amati Global Investors |
572,459 |
£9,044,852 |
| Chelverton Asset Management |
569,750 |
£9,002,050 |
| Brooks Macdonald Employee Benefit Trust |
552,633 |
£8,731,601 |
| Rathbones Investment Management |
488,191 |
£7,713,418 |
| Kames Capital |
487,376 |
£7,700,541 |
| Spearpoint Holdings Limited |
428,227 |
£6,765,987 |