| Gresham House Asset Management Ltd |
3,746,521 |
£59,944,336 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£52,457,632 |
| Liontrust Investment Services Limited |
2,457,011 |
£39,312,176 |
| Octopus Investments Nominees Limited |
2,335,294 |
£37,364,704 |
| Brooks Macdonald Asset Management |
1,105,946 |
£17,695,136 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,340,608 |
| Jupiter Fund Management plc |
938,000 |
£15,008,000 |
| Aberforth Partners LLP |
898,030 |
£14,368,480 |
| Invesco Asset Management |
817,151 |
£13,074,416 |
| Invesco |
817,151 |
£13,074,416 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£12,947,200 |
| The Wellcome Trust Limited |
805,370 |
£12,885,920 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,583,232 |
| Hargreave Hale & Co |
784,942 |
£12,559,072 |
| Canaccord Genuity Group Inc |
780,008 |
£12,480,128 |
| Artemis Investment Management Ltd |
729,575 |
£11,673,200 |
| Norges Bank |
683,030 |
£10,928,480 |
| Charles Stanley |
659,277 |
£10,548,432 |
| Jonathan M Gumpel |
642,460 |
£10,279,360 |
| Brewin Dolphin Limited |
622,157 |
£9,954,512 |
| JTC plc |
603,261 |
£9,652,176 |
| Amati Global Investors |
572,459 |
£9,159,344 |
| Chelverton Asset Management |
569,750 |
£9,116,000 |
| Brooks Macdonald Employee Benefit Trust |
552,633 |
£8,842,128 |
| Rathbones Investment Management |
488,191 |
£7,811,056 |
| Kames Capital |
487,376 |
£7,798,016 |
| Spearpoint Holdings Limited |
428,227 |
£6,851,632 |