| Alantra EQMC Asset Management SGIIC, S.A. |
4,876,250 |
£81,140,800 |
| EQMC Europe Development Capital Fund |
4,858,350 |
£80,842,944 |
| Kempen Capital Management N.V. |
3,328,822 |
£55,391,598 |
| BlackRock Smaller Companies Trust Plc |
3,206,055 |
£53,348,755 |
| Capital Research and Management Company |
2,990,645 |
£49,764,333 |
| Aberdeen Asset Management plc |
1,835,099 |
£30,536,047 |
| Franklin Templeton Investments |
1,700,000 |
£28,288,000 |
| Threadneedle Asset Management |
1,597,700 |
£26,585,728 |
| Aggregate of Standard Life Aberdeen plc |
1,580,030 |
£26,291,699 |
| River & Mercantile Asset Management LLP |
1,549,478 |
£25,783,314 |
| Hargreave Hale & Co |
1,546,771 |
£25,738,269 |
| Cazenove Capital Management Limited |
1,538,403 |
£25,599,026 |
| Ameriprise Financial, Inc. and its group |
1,538,156 |
£25,594,916 |
| Canaccord Genuity Group Inc |
1,529,075 |
£25,443,808 |
| Polar Capital LLP |
1,517,909 |
£25,258,006 |
| Ancora Alternatives LLC |
1,504,769 |
£25,039,356 |
| MI Chelverton UK Equity Growth Fund |
1,500,000 |
£24,960,000 |
| ClearBridge Investment Management Limited |
1,486,511 |
£24,735,543 |
| Schroders plc |
1,478,291 |
£24,598,762 |
| The Capital Group Companies, Inc. |
1,443,660 |
£24,022,502 |
| Aberforth Partners LLP |
1,416,081 |
£23,563,588 |
| Janus Henderson Investors |
1,408,457 |
£23,436,724 |
| Wasatch Advisors, Inc. |
1,285,950 |
£21,398,208 |
| JPMorgan Asset Management Holdings Inc. |
1,256,443 |
£20,907,212 |
| JTC Share Plan Trustees (Guernsey) Limited |
878,463 |
£14,617,624 |