| Alantra EQMC Asset Management SGIIC, S.A. |
4,876,250 |
£84,066,550 |
| EQMC Europe Development Capital Fund |
4,858,350 |
£83,757,954 |
| Kempen Capital Management N.V. |
3,328,822 |
£57,388,891 |
| BlackRock Smaller Companies Trust Plc |
3,206,055 |
£55,272,388 |
| Capital Research and Management Company |
2,990,645 |
£51,558,720 |
| Aberdeen Asset Management plc |
1,835,099 |
£31,637,107 |
| Franklin Templeton Investments |
1,700,000 |
£29,308,000 |
| Threadneedle Asset Management |
1,597,700 |
£27,544,348 |
| Aggregate of Standard Life Aberdeen plc |
1,580,030 |
£27,239,717 |
| River & Mercantile Asset Management LLP |
1,549,478 |
£26,713,001 |
| Hargreave Hale & Co |
1,546,771 |
£26,666,332 |
| Cazenove Capital Management Limited |
1,538,403 |
£26,522,068 |
| Ameriprise Financial, Inc. and its group |
1,538,156 |
£26,517,809 |
| Canaccord Genuity Group Inc |
1,529,075 |
£26,361,253 |
| Polar Capital LLP |
1,517,909 |
£26,168,751 |
| Ancora Alternatives LLC |
1,504,769 |
£25,942,218 |
| MI Chelverton UK Equity Growth Fund |
1,500,000 |
£25,860,000 |
| ClearBridge Investment Management Limited |
1,486,511 |
£25,627,450 |
| Schroders plc |
1,478,291 |
£25,485,737 |
| The Capital Group Companies, Inc. |
1,443,660 |
£24,888,698 |
| BlackRock, Inc. |
1,443,454 |
£24,885,147 |
| Aberforth Partners LLP |
1,416,081 |
£24,413,236 |
| Janus Henderson Investors |
1,408,457 |
£24,281,799 |
| Wasatch Advisors, Inc. |
1,285,950 |
£22,169,778 |
| JPMorgan Asset Management Holdings Inc. |
1,256,443 |
£21,661,077 |
| JTC Share Plan Trustees (Guernsey) Limited |
878,463 |
£15,144,702 |