| Gresham House Asset Management Ltd |
3,746,521 |
£62,754,227 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£54,916,584 |
| Liontrust Investment Services Limited |
2,348,411 |
£39,335,884 |
| Octopus Investments Nominees Limited |
2,335,294 |
£39,116,174 |
| Aberforth Partners LLP |
1,606,852 |
£26,914,771 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£18,626,251 |
| Brooks Macdonald Asset Management |
1,105,946 |
£18,524,596 |
| Canaccord Genuity Group Inc |
1,021,288 |
£17,106,574 |
| Jupiter Fund Management plc |
938,000 |
£15,711,500 |
| Invesco Asset Management |
817,151 |
£13,687,279 |
| Invesco |
817,151 |
£13,687,279 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£13,554,100 |
| The Wellcome Trust Limited |
805,370 |
£13,489,948 |
| FIDELITY INTERNATIONAL |
786,452 |
£13,173,071 |
| Hargreave Hale & Co |
784,942 |
£13,147,778 |
| Canaccord Genuity Group Inc |
780,008 |
£13,065,134 |
| Artemis Investment Management Ltd |
729,575 |
£12,220,381 |
| Norges Bank |
683,030 |
£11,440,752 |
| Charles Stanley |
659,277 |
£11,042,890 |
| Jonathan M Gumpel |
642,460 |
£10,761,205 |
| Brewin Dolphin Limited |
622,157 |
£10,421,130 |
| Chelverton Asset Management |
609,008 |
£10,200,884 |
| JTC plc |
603,261 |
£10,104,622 |
| Amati Global Investors |
572,459 |
£9,588,688 |
| Rathbones Investment Management |
488,191 |
£8,177,199 |
| Kames Capital |
487,376 |
£8,163,548 |
| Spearpoint Holdings Limited |
428,227 |
£7,172,802 |