| Gresham House Asset Management Ltd |
3,746,521 |
£63,690,857 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£55,736,234 |
| Liontrust Investment Services Limited |
2,457,011 |
£41,769,187 |
| Octopus Investments Nominees Limited |
2,335,294 |
£39,699,998 |
| Aberforth Partners LLP |
1,606,852 |
£27,316,484 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£18,904,255 |
| Brooks Macdonald Asset Management |
1,105,946 |
£18,801,082 |
| Canaccord Genuity Group Inc |
1,021,288 |
£17,361,896 |
| Jupiter Fund Management plc |
938,000 |
£15,946,000 |
| Invesco Asset Management |
817,151 |
£13,891,567 |
| Invesco |
817,151 |
£13,891,567 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£13,756,400 |
| The Wellcome Trust Limited |
805,370 |
£13,691,290 |
| FIDELITY INTERNATIONAL |
786,452 |
£13,369,684 |
| Hargreave Hale & Co |
784,942 |
£13,344,014 |
| Canaccord Genuity Group Inc |
780,008 |
£13,260,136 |
| Artemis Investment Management Ltd |
729,575 |
£12,402,775 |
| Norges Bank |
683,030 |
£11,611,510 |
| Charles Stanley |
659,277 |
£11,207,709 |
| Jonathan M Gumpel |
642,460 |
£10,921,820 |
| Brewin Dolphin Limited |
622,157 |
£10,576,669 |
| Chelverton Asset Management |
609,008 |
£10,353,136 |
| JTC plc |
603,261 |
£10,255,437 |
| Amati Global Investors |
572,459 |
£9,731,803 |
| Rathbones Investment Management |
488,191 |
£8,299,247 |
| Kames Capital |
487,376 |
£8,285,392 |
| Spearpoint Holdings Limited |
428,227 |
£7,279,859 |