| Gresham House Asset Management Ltd |
3,746,521 |
£60,693,640 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£53,113,352 |
| Liontrust Investment Services Limited |
2,457,011 |
£39,803,578 |
| Octopus Investments Nominees Limited |
2,335,294 |
£37,831,763 |
| Aberforth Partners LLP |
2,067,205 |
£33,488,721 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£18,014,643 |
| Brooks Macdonald Asset Management |
1,105,946 |
£17,916,325 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,544,866 |
| Jupiter Fund Management plc |
938,000 |
£15,195,600 |
| Invesco Asset Management |
817,151 |
£13,237,846 |
| Invesco |
817,151 |
£13,237,846 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£13,109,040 |
| The Wellcome Trust Limited |
805,370 |
£13,046,994 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,740,522 |
| Hargreave Hale & Co |
784,942 |
£12,716,060 |
| Canaccord Genuity Group Inc |
780,008 |
£12,636,130 |
| Artemis Investment Management Ltd |
729,575 |
£11,819,115 |
| Norges Bank |
683,030 |
£11,065,086 |
| Charles Stanley |
659,277 |
£10,680,287 |
| Jonathan M Gumpel |
642,460 |
£10,407,852 |
| Brewin Dolphin Limited |
622,157 |
£10,078,943 |
| Chelverton Asset Management |
609,008 |
£9,865,930 |
| JTC plc |
603,261 |
£9,772,828 |
| Amati Global Investors |
572,459 |
£9,273,836 |
| Rathbones Investment Management |
488,191 |
£7,908,694 |
| Kames Capital |
487,376 |
£7,895,491 |
| Spearpoint Holdings Limited |
428,227 |
£6,937,277 |