| Andrew William Oliver |
205,882,352,941 |
£360,294,118 |
| TD Direct Investing Nominees |
94,495,144,706 |
£165,366,503 |
| HSDL Nominees Limited |
81,968,497,647 |
£143,444,871 |
| David Lenigas |
79,411,764,706 |
£138,970,588 |
| Greenhair Services Limited |
73,235,294,118 |
£128,161,765 |
| Academy Minerals Limited |
70,588,235,294 |
£123,529,412 |
| Kavi Dhana |
44,021,067,059 |
£77,036,867 |
| IP2IPO Limited (together with Techtran Group Limited) |
34,418,058,824 |
£60,231,603 |
| XCAP Nominees Limited |
19,824,847,059 |
£34,693,482 |
| IP Venture Fund Limited |
16,302,105,882 |
£28,528,685 |
| Mr Mark V Houghton-Berry |
12,335,376,471 |
£21,586,909 |
| The University of Leeds |
9,552,264,706 |
£16,716,463 |
| The Wellcome Trust Limited |
6,107,858,824 |
£10,688,753 |
| Adrian Stiff |
37,000,000 |
£64,750 |
| Hargreaves Lansdown |
27,752,212 |
£48,566 |
| JIM Nominees Limited |
27,116,707 |
£47,454 |
| Pershing Nominees |
26,006,789 |
£45,512 |
| Hargreaves Lansdown (Nominees) Limited Des 15942 |
25,609,943 |
£44,817 |
| Chris Akers |
25,000,000 |
£43,750 |
| Vidacos Nominees Limited |
20,709,150 |
£36,241 |
| The Bank OF New York (Nominees) Limited Des 672938 |
19,705,371 |
£34,484 |
| Oberon Investments |
19,705,371 |
£34,484 |
| Barclays Direct Investing Nominees Limited |
19,358,393 |
£33,877 |
| Winterflood Securities Limited |
18,278,266 |
£31,987 |
| Donald Strang |
17,820,211 |
£31,185 |
| Interactive Investor Services Nominees Limited |
17,219,649 |
£30,134 |
| ABN Amro Bank |
13,801,956 |
£24,153 |
| Hargreaves Lansdown (Nominees) Limited Des VRA |
12,572,701 |
£22,002 |
| David Graham Brown |
12,100,000 |
£21,175 |
| Sebastian Marr |
8,180,672 |
£14,316 |