| Gresham House Asset Management Ltd |
3,746,521 |
£60,318,988 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£52,785,492 |
| Liontrust Investment Services Limited |
2,457,011 |
£39,557,877 |
| Octopus Investments Nominees Limited |
2,335,294 |
£37,598,233 |
| Aberforth Partners LLP |
2,067,205 |
£33,282,000 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£17,903,442 |
| Brooks Macdonald Asset Management |
1,105,946 |
£17,805,731 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,442,737 |
| Jupiter Fund Management plc |
938,000 |
£15,101,800 |
| Invesco Asset Management |
817,151 |
£13,156,131 |
| Invesco |
817,151 |
£13,156,131 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£13,028,120 |
| The Wellcome Trust Limited |
805,370 |
£12,966,457 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,661,877 |
| Hargreave Hale & Co |
784,942 |
£12,637,566 |
| Canaccord Genuity Group Inc |
780,008 |
£12,558,129 |
| Artemis Investment Management Ltd |
729,575 |
£11,746,158 |
| Norges Bank |
683,030 |
£10,996,783 |
| Charles Stanley |
659,277 |
£10,614,360 |
| Jonathan M Gumpel |
642,460 |
£10,343,606 |
| Brewin Dolphin Limited |
622,157 |
£10,016,728 |
| Chelverton Asset Management |
609,008 |
£9,805,029 |
| JTC plc |
603,261 |
£9,712,502 |
| Amati Global Investors |
572,459 |
£9,216,590 |
| Rathbones Investment Management |
488,191 |
£7,859,875 |
| Kames Capital |
487,376 |
£7,846,754 |
| Spearpoint Holdings Limited |
428,227 |
£6,894,455 |