| Alantra EQMC Asset Management SGIIC, S.A. |
4,876,250 |
£89,527,950 |
| EQMC Europe Development Capital Fund |
4,858,350 |
£89,199,306 |
| Kempen Capital Management N.V. |
3,328,822 |
£61,117,172 |
| BlackRock Smaller Companies Trust Plc |
3,206,055 |
£58,863,170 |
| Capital Research and Management Company |
2,990,645 |
£54,908,242 |
| Aberdeen Asset Management plc |
1,835,099 |
£33,692,418 |
| Franklin Templeton Investments |
1,700,000 |
£31,212,000 |
| Threadneedle Asset Management |
1,597,700 |
£29,333,772 |
| Aggregate of Standard Life Aberdeen plc |
1,580,030 |
£29,009,351 |
| River & Mercantile Asset Management LLP |
1,549,478 |
£28,448,416 |
| Hargreave Hale & Co |
1,546,771 |
£28,398,716 |
| Cazenove Capital Management Limited |
1,538,403 |
£28,245,079 |
| Ameriprise Financial, Inc. and its group |
1,538,156 |
£28,240,544 |
| Canaccord Genuity Group Inc |
1,529,075 |
£28,073,817 |
| Polar Capital LLP |
1,517,909 |
£27,868,809 |
| Ancora Alternatives LLC |
1,504,769 |
£27,627,559 |
| MI Chelverton UK Equity Growth Fund |
1,500,000 |
£27,540,000 |
| ClearBridge Investment Management Limited |
1,486,511 |
£27,292,342 |
| Schroders plc |
1,478,291 |
£27,141,423 |
| The Capital Group Companies, Inc. |
1,443,660 |
£26,505,598 |
| BlackRock, Inc. |
1,443,454 |
£26,501,815 |
| Aberforth Partners LLP |
1,416,081 |
£25,999,247 |
| Janus Henderson Investors |
1,408,457 |
£25,859,270 |
| Wasatch Advisors, Inc. |
1,285,950 |
£23,610,042 |
| JPMorgan Asset Management Holdings Inc. |
1,256,443 |
£23,068,294 |
| JTC Share Plan Trustees (Guernsey) Limited |
878,463 |
£16,128,581 |