| Alantra EQMC Asset Management SGIIC, S.A. |
4,876,250 |
£90,503,200 |
| EQMC Europe Development Capital Fund |
4,858,350 |
£90,170,976 |
| Kempen Capital Management N.V. |
3,328,822 |
£61,782,936 |
| BlackRock Smaller Companies Trust Plc |
3,206,055 |
£59,504,381 |
| Capital Research and Management Company |
2,990,645 |
£55,506,371 |
| Aberdeen Asset Management plc |
1,835,099 |
£34,059,437 |
| Franklin Templeton Investments |
1,700,000 |
£31,552,000 |
| Threadneedle Asset Management |
1,597,700 |
£29,653,312 |
| Aggregate of Standard Life Aberdeen plc |
1,580,030 |
£29,325,357 |
| River & Mercantile Asset Management LLP |
1,549,478 |
£28,758,312 |
| Hargreave Hale & Co |
1,546,771 |
£28,708,070 |
| Cazenove Capital Management Limited |
1,538,403 |
£28,552,760 |
| Ameriprise Financial, Inc. and its group |
1,538,156 |
£28,548,175 |
| Canaccord Genuity Group Inc |
1,529,075 |
£28,379,632 |
| Polar Capital LLP |
1,517,909 |
£28,172,391 |
| Ancora Alternatives LLC |
1,504,769 |
£27,928,513 |
| MI Chelverton UK Equity Growth Fund |
1,500,000 |
£27,840,000 |
| ClearBridge Investment Management Limited |
1,486,511 |
£27,589,644 |
| Schroders plc |
1,478,291 |
£27,437,081 |
| The Capital Group Companies, Inc. |
1,443,660 |
£26,794,330 |
| BlackRock, Inc. |
1,443,454 |
£26,790,506 |
| Aberforth Partners LLP |
1,416,081 |
£26,282,463 |
| Janus Henderson Investors |
1,408,457 |
£26,140,962 |
| Wasatch Advisors, Inc. |
1,285,950 |
£23,867,232 |
| JPMorgan Asset Management Holdings Inc. |
1,256,443 |
£23,319,582 |
| JTC Share Plan Trustees (Guernsey) Limited |
878,463 |
£16,304,273 |