| Gresham House Asset Management Ltd |
3,746,521 |
£61,630,270 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£53,933,003 |
| Liontrust Investment Services Limited |
2,457,011 |
£40,417,831 |
| Octopus Investments Nominees Limited |
2,335,294 |
£38,415,586 |
| Brooks Macdonald Asset Management |
1,105,946 |
£18,192,812 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,800,188 |
| Jupiter Fund Management plc |
938,000 |
£15,430,100 |
| Aberforth Partners LLP |
898,030 |
£14,772,594 |
| Invesco Asset Management |
817,151 |
£13,442,134 |
| Invesco |
817,151 |
£13,442,134 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£13,311,340 |
| The Wellcome Trust Limited |
805,370 |
£13,248,336 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,937,135 |
| Hargreave Hale & Co |
784,942 |
£12,912,296 |
| Canaccord Genuity Group Inc |
780,008 |
£12,831,132 |
| Artemis Investment Management Ltd |
729,575 |
£12,001,509 |
| Norges Bank |
683,030 |
£11,235,844 |
| Charles Stanley |
659,277 |
£10,845,107 |
| Jonathan M Gumpel |
642,460 |
£10,568,467 |
| Brewin Dolphin Limited |
622,157 |
£10,234,483 |
| JTC plc |
603,261 |
£9,923,644 |
| Amati Global Investors |
572,459 |
£9,416,950 |
| Chelverton Asset Management |
569,750 |
£9,372,388 |
| Brooks Macdonald Employee Benefit Trust |
552,633 |
£9,090,813 |
| Rathbones Investment Management |
488,191 |
£8,030,742 |
| Kames Capital |
487,376 |
£8,017,335 |
| Spearpoint Holdings Limited |
428,227 |
£7,044,334 |