| Andrew William Oliver |
205,882,352,941 |
£240,196,765 |
| TD Direct Investing Nominees |
94,495,144,706 |
£110,244,650 |
| HSDL Nominees Limited |
81,968,497,647 |
£95,630,187 |
| David Lenigas |
79,411,764,706 |
£92,647,324 |
| Greenhair Services Limited |
73,235,294,118 |
£85,441,421 |
| Academy Minerals Limited |
70,588,235,294 |
£82,353,176 |
| Kavi Dhana |
44,021,067,059 |
£51,358,058 |
| IP2IPO Limited (together with Techtran Group Limited) |
34,418,058,824 |
£40,154,517 |
| XCAP Nominees Limited |
19,824,847,059 |
£23,129,054 |
| IP Venture Fund Limited |
16,302,105,882 |
£19,019,178 |
| Mr Mark V Houghton-Berry |
12,335,376,471 |
£14,391,314 |
| The University of Leeds |
9,552,264,706 |
£11,144,341 |
| The Wellcome Trust Limited |
6,107,858,824 |
£7,125,856 |
| Adrian Stiff |
37,000,000 |
£43,167 |
| Hargreaves Lansdown |
27,752,212 |
£32,378 |
| JIM Nominees Limited |
27,116,707 |
£31,636 |
| Pershing Nominees |
26,006,789 |
£30,341 |
| Hargreaves Lansdown (Nominees) Limited Des 15942 |
25,609,943 |
£29,878 |
| Chris Akers |
25,000,000 |
£29,167 |
| Vidacos Nominees Limited |
20,709,150 |
£24,161 |
| The Bank OF New York (Nominees) Limited Des 672938 |
19,705,371 |
£22,990 |
| Oberon Investments |
19,705,371 |
£22,990 |
| Barclays Direct Investing Nominees Limited |
19,358,393 |
£22,585 |
| Winterflood Securities Limited |
18,278,266 |
£21,325 |
| Donald Strang |
17,820,211 |
£20,790 |
| Interactive Investor Services Nominees Limited |
17,219,649 |
£20,090 |
| ABN Amro Bank |
13,801,956 |
£16,102 |
| Hargreaves Lansdown (Nominees) Limited Des VRA |
12,572,701 |
£14,668 |
| David Graham Brown |
12,100,000 |
£14,117 |
| Sebastian Marr |
8,180,672 |
£9,544 |