| Gresham House Asset Management Ltd |
3,746,521 |
£61,442,944 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£53,769,073 |
| Liontrust Investment Services Limited |
2,457,011 |
£40,294,980 |
| Octopus Investments Nominees Limited |
2,335,294 |
£38,298,822 |
| Brooks Macdonald Asset Management |
1,105,946 |
£18,137,514 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,749,123 |
| Jupiter Fund Management plc |
938,000 |
£15,383,200 |
| Aberforth Partners LLP |
898,030 |
£14,727,692 |
| Invesco Asset Management |
817,151 |
£13,401,276 |
| Invesco |
817,151 |
£13,401,276 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£13,270,880 |
| The Wellcome Trust Limited |
805,370 |
£13,208,068 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,897,813 |
| Hargreave Hale & Co |
784,942 |
£12,873,049 |
| Canaccord Genuity Group Inc |
780,008 |
£12,792,131 |
| Artemis Investment Management Ltd |
729,575 |
£11,965,030 |
| Norges Bank |
683,030 |
£11,201,692 |
| Charles Stanley |
659,277 |
£10,812,143 |
| Jonathan M Gumpel |
642,460 |
£10,536,344 |
| Brewin Dolphin Limited |
622,157 |
£10,203,375 |
| JTC plc |
603,261 |
£9,893,480 |
| Amati Global Investors |
572,459 |
£9,388,328 |
| Chelverton Asset Management |
569,750 |
£9,343,900 |
| Brooks Macdonald Employee Benefit Trust |
552,633 |
£9,063,181 |
| Rathbones Investment Management |
488,191 |
£8,006,332 |
| Kames Capital |
487,376 |
£7,992,966 |
| Spearpoint Holdings Limited |
428,227 |
£7,022,923 |