| Alantra EQMC Asset Management SGIIC, S.A. |
4,876,250 |
£94,989,350 |
| EQMC Europe Development Capital Fund |
4,858,350 |
£94,640,658 |
| Kempen Capital Management N.V. |
3,328,822 |
£64,845,453 |
| BlackRock Smaller Companies Trust Plc |
3,206,055 |
£62,453,951 |
| Capital Research and Management Company |
2,990,645 |
£58,257,765 |
| Aberdeen Asset Management plc |
1,835,099 |
£35,747,728 |
| Franklin Templeton Investments |
1,700,000 |
£33,116,000 |
| Threadneedle Asset Management |
1,597,700 |
£31,123,196 |
| Aggregate of Standard Life Aberdeen plc |
1,580,030 |
£30,778,984 |
| River & Mercantile Asset Management LLP |
1,549,478 |
£30,183,831 |
| Hargreave Hale & Co |
1,546,771 |
£30,131,099 |
| Cazenove Capital Management Limited |
1,538,403 |
£29,968,090 |
| Ameriprise Financial, Inc. and its group |
1,538,156 |
£29,963,279 |
| Canaccord Genuity Group Inc |
1,529,075 |
£29,786,381 |
| Polar Capital LLP |
1,517,909 |
£29,568,867 |
| Ancora Alternatives LLC |
1,504,769 |
£29,312,900 |
| MI Chelverton UK Equity Growth Fund |
1,500,000 |
£29,220,000 |
| ClearBridge Investment Management Limited |
1,486,511 |
£28,957,234 |
| Schroders plc |
1,478,291 |
£28,797,109 |
| The Capital Group Companies, Inc. |
1,443,660 |
£28,122,497 |
| BlackRock, Inc. |
1,443,454 |
£28,118,484 |
| Aberforth Partners LLP |
1,416,081 |
£27,585,258 |
| Janus Henderson Investors |
1,408,457 |
£27,436,742 |
| Wasatch Advisors, Inc. |
1,285,950 |
£25,050,306 |
| JPMorgan Asset Management Holdings Inc. |
1,256,443 |
£24,475,510 |
| JTC Share Plan Trustees (Guernsey) Limited |
878,463 |
£17,112,459 |