| Gresham House Asset Management Ltd |
3,746,521 |
£62,192,249 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£54,424,793 |
| Liontrust Investment Services Limited |
2,457,011 |
£40,786,383 |
| Octopus Investments Nominees Limited |
2,335,294 |
£38,765,880 |
| Aberforth Partners LLP |
2,067,205 |
£34,315,603 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£18,459,449 |
| Brooks Macdonald Asset Management |
1,105,946 |
£18,358,704 |
| Canaccord Genuity Group Inc |
1,021,288 |
£16,953,381 |
| Jupiter Fund Management plc |
938,000 |
£15,570,800 |
| Invesco Asset Management |
817,151 |
£13,564,707 |
| Invesco |
817,151 |
£13,564,707 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£13,432,720 |
| The Wellcome Trust Limited |
805,370 |
£13,369,142 |
| FIDELITY INTERNATIONAL |
786,452 |
£13,055,103 |
| Hargreave Hale & Co |
784,942 |
£13,030,037 |
| Canaccord Genuity Group Inc |
780,008 |
£12,948,133 |
| Artemis Investment Management Ltd |
729,575 |
£12,110,945 |
| Norges Bank |
683,030 |
£11,338,298 |
| Charles Stanley |
659,277 |
£10,943,998 |
| Jonathan M Gumpel |
642,460 |
£10,664,836 |
| Brewin Dolphin Limited |
622,157 |
£10,327,806 |
| Chelverton Asset Management |
609,008 |
£10,109,533 |
| JTC plc |
603,261 |
£10,014,133 |
| Amati Global Investors |
572,459 |
£9,502,819 |
| Rathbones Investment Management |
488,191 |
£8,103,971 |
| Kames Capital |
487,376 |
£8,090,442 |
| Spearpoint Holdings Limited |
428,227 |
£7,108,568 |