| MassMutual Life Insurance Company |
12,727,105 |
£523,847,642 |
| MFS Investment Management |
11,499,077 |
£473,302,009 |
| Fidelity Management & Research Company |
8,063,819 |
£331,906,790 |
| MI Metron UK Bidco Limited |
7,691,714 |
£316,590,948 |
| Societe Generale |
7,177,114 |
£295,410,012 |
| Standard Life Investment Ltd |
5,920,593 |
£243,691,608 |
| OppenheimerFunds, Inc. |
5,720,217 |
£235,444,132 |
| Marathon Asset Management LLP |
5,708,763 |
£234,972,685 |
| Vanguard Group |
5,394,997 |
£222,058,076 |
| UBS Asset Management |
5,231,636 |
£215,334,138 |
| F & C Asset Management plc |
5,212,550 |
£214,548,558 |
| Massachusetts Financial Services Company |
5,178,500 |
£213,147,060 |
| JPMorgan Chase & Co. |
4,841,669 |
£199,283,096 |
| Wellington Management Company |
4,248,835 |
£174,882,049 |
| Sprucegrove Investment Management |
3,832,503 |
£157,745,824 |
| Artemis Investment Management |
3,661,077 |
£150,689,929 |
| Royal London Asset Management Limited |
3,626,169 |
£149,253,116 |
| Schroder Investment Management |
3,484,265 |
£143,412,347 |
| BlackRock, Inc. |
3,429,918 |
£141,175,425 |
| Ninety One |
3,160,918 |
£130,103,385 |
| Liontrust Asset Management |
3,052,149 |
£125,626,453 |
| Evenlode Investment |
2,971,067 |
£122,289,118 |
| Ignis Investment Services Limited |
10 |
£412 |
| Ameriprise Financial Inc. |
1 |
£41 |
| COLUMBIA WANGER ASSET MANAGEMENT, L.P. |
1 |
£41 |