| Gresham House Asset Management Ltd |
3,884,684 |
£56,327,918 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£47,539,729 |
| Octopus Investments Nominees Limited |
2,335,294 |
£33,861,763 |
| Liontrust Investment Services Limited |
2,215,411 |
£32,123,460 |
| Aberforth Partners LLP |
1,606,852 |
£23,299,354 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£16,124,218 |
| Brooks Macdonald Asset Management |
1,105,946 |
£16,036,217 |
| Canaccord Genuity Group Inc |
1,021,288 |
£14,808,676 |
| Jupiter Fund Management plc |
938,000 |
£13,601,000 |
| Invesco Asset Management |
817,151 |
£11,848,690 |
| Invesco |
817,151 |
£11,848,690 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£11,733,400 |
| The Wellcome Trust Limited |
805,370 |
£11,677,865 |
| FIDELITY INTERNATIONAL |
786,452 |
£11,403,554 |
| Hargreave Hale & Co |
784,942 |
£11,381,659 |
| Canaccord Genuity Group Inc |
780,008 |
£11,310,116 |
| Artemis Investment Management Ltd |
729,575 |
£10,578,838 |
| Norges Bank |
683,030 |
£9,903,935 |
| Charles Stanley |
659,277 |
£9,559,516 |
| Jonathan M Gumpel |
642,460 |
£9,315,670 |
| Brewin Dolphin Limited |
622,157 |
£9,021,276 |
| Chelverton Asset Management |
609,008 |
£8,830,616 |
| JTC plc |
603,261 |
£8,747,284 |
| Amati Global Investors |
572,459 |
£8,300,656 |
| Rathbones Investment Management |
488,191 |
£7,078,770 |
| Kames Capital |
487,376 |
£7,066,952 |
| Spearpoint Holdings Limited |
428,227 |
£6,209,292 |