| Gresham House Asset Management Ltd |
3,746,521 |
£58,445,728 |
| Montanaro UK Smaller Companies Investment Trust plc |
3,278,602 |
£51,146,191 |
| Liontrust Investment Services Limited |
2,348,411 |
£36,635,212 |
| Octopus Investments Nominees Limited |
2,335,294 |
£36,430,586 |
| Aberforth Partners LLP |
1,606,852 |
£25,066,891 |
| Brooks Macdonald Employee Benefit Trust |
1,112,015 |
£17,347,434 |
| Brooks Macdonald Asset Management |
1,105,946 |
£17,252,758 |
| Canaccord Genuity Group Inc |
1,021,288 |
£15,932,093 |
| Jupiter Fund Management plc |
938,000 |
£14,632,800 |
| Invesco Asset Management |
817,151 |
£12,747,556 |
| Invesco |
817,151 |
£12,747,556 |
| MI Chelverton UK Equity Growth Fund |
809,200 |
£12,623,520 |
| The Wellcome Trust Limited |
805,370 |
£12,563,772 |
| FIDELITY INTERNATIONAL |
786,452 |
£12,268,651 |
| Hargreave Hale & Co |
784,942 |
£12,245,095 |
| Canaccord Genuity Group Inc |
780,008 |
£12,168,125 |
| Artemis Investment Management Ltd |
729,575 |
£11,381,370 |
| Norges Bank |
683,030 |
£10,655,268 |
| Charles Stanley |
659,277 |
£10,284,721 |
| Jonathan M Gumpel |
642,460 |
£10,022,376 |
| Brewin Dolphin Limited |
622,157 |
£9,705,649 |
| Chelverton Asset Management |
609,008 |
£9,500,525 |
| JTC plc |
603,261 |
£9,410,872 |
| Amati Global Investors |
572,459 |
£8,930,360 |
| Rathbones Investment Management |
488,191 |
£7,615,780 |
| Kames Capital |
487,376 |
£7,603,066 |
| Spearpoint Holdings Limited |
428,227 |
£6,680,341 |